The line of scrimmage is where the football is placed at the beginning of each play. This line can move forward or backward depending on the gain or loss of each play.
Scrimmage, derived from an obsolete variant of skirmish. It basically means a rough-and-tumble struggle; a skirmish. In American football it refers to the contest of each play, beginning with the snap and ending with the dead ball whistle; or a team's practice session.
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In football the position behind the line of scrimmage changes depending on if the offense, defense, or special teams is on the field. In the offense it is the quarterback and possibly the running backs. The defense has the safeties, linebackers, and defensive backs playing off the line. In special teams it would be the kicker or punter.