To be able to swim fast, but preferable to be able to accelerate very fast and to start faster than your opponent. Next you need a lot of stamina and you need to be able to tread water using the eggbeater technique. This involves kicking your legs one after the other instead of kicking them simultaneously. This will make sure you don't need your arms to stay above water, so you can use them to throw the ball or block your opponents' throw.
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in order to be a decent swimmer you have to know at least backstroke and freestyle well. that means you can do it with out getting super tired.
you need to be able to throw the ball far, stay above water for a long time, swim fast, have fast reflexes, be able to catch the ball, and you need to be able to gage your shot.
Three most important Water Polo skills are Treadingwater,Swimming and Ball handling skills.