ANSWER:For the front wheel on most bikes, it wont matter. On The one I had, the rear axle had a specific groove for where the sprocket went and the axle wasnt interchangeable. You knew right away if It was on backwards because the chain wouldn't go on the bike. For the newer bikes, I truly don't know.
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More answers
You can change the axle length by installing a longer axle. You
cannot change the axle diameter because you are limited to the
diameter of the bearing cups, races, etc. allowed by whatever
particular hub is being used.